Ram Singh
Agronomist
236 National Soybean Res Ctr 103 S. Goodwin
Urbana, IL 61801
(217) 333-8144
Primary Disciplines
Education
Ph.D.: Genetics, Colorado State University, Fort Collins, Colorado — 1974
M.S.: Agricultural Botany, passed in the first division, Government Agricultural College, Kanpur, Uttar Pradesh, India — 1968
B.S.: Agriculture, passed in the first division, Government Agricultural College, Kanpur, Uttar Pradesh, India — 1966
Intermediate: Passed in the first division with distinction in Agricultural Engineering and Agronomy, Edai Pratap Inter College, Varanasi, U.P Varanasi, Uttar Pradesh, India — 1963
High School: Passed in first division with distinction in Sanskrit, Kalika Dham Inter College, Sewapuri, Varanasi, Uttar Pradesh, India 1961
Research Areas
Germplasm Maintenance, Plant Cytogenetics, Plant Genome Mapping, Genomics, Plant Breeding, Biosystematics, Wide Hybridization, Cell and Tissue Culture, Plant Genetic Transformation, Plant Molecular Cytogenetics
Positions
Agronomist, Soybean Cytogenetics, University of Illinois, Urbana-Champaign, Illinois (1986 - present)
Associate Agronomist, Soybean Cytogenetics, University of Illinois, Urbana-Champaign, Illinois (1983 - 1986)
Research Scientist, Barley and Potato Cytogenetics, Cell and Tissue Culture, Advanced Genetic Sciences, Inc., Manhattan, Kansas (1981 - 1982)
Visiting Scientist, Barley Cytogenetics, Colorado State University, Fort Collins, Colorado (1980 - 1981)
Senior Research Fellow, Rice Cytogenetics and Breeding, International Rice Research Institute, Philippines (1977 - 1980)
Visiting Scientist, Wheat and Rye Cytogenetics, University of Göttingen, Göttingen, Germany (1974 - 1976)
Research Assistant, Barley Cytogenetics, Colorado State University, Fort Collins, Colorado (1971 - 1974)
Research Assistant, Germplasm Maintenance and Seed Testing, The Rockefeller Foundation, New Delhi, India (1968 - 1970)
Honors and Awards
Foreign Fellow, National Academy of Agricultural Sciences (NAAS), India, 2012
The 2010 Illinois Soybean Association’s Excellence in Soybean Research Award
College of ACES Professional Staff Award for Excellence – Research, 2009
Academic Professional Award for Excellence 2007
Academic Professional Award for Excellence: Innovative & Creativity, 2000
The Marquis Who'sWho in America 55th edition, 2001
Senior Research Fellow, International Rice Research Institute, Las Baños, Philippines, 1977-1980
First Year University Fellow, University of California, Davis, California, 1970-1971
Senior Research Fellow, Indian Council Agriculture Research, New Delhi, 1966-1968
Journal Articles
Chung, G., and R. J. Singh. 2008. Broadening the genetic base of soybean: a multi-disciplinary approach. Critical Rev. Plant Sci. 27: 295-341.
Singh, R. J. and G. H. Chung. 2007. Cytogenetics of Soybean: Progress and Prospective. The Nucleus 50: 401-423.
Singh, R. J., G. H. Chung, and R. L. Nelson. 2007. Landmark research in legumes. Genome 50: 525-537.
Zou, J. J., R. J. Singh, J. Lee, S. J. Xu and T. Hymowitz. 2006. SSR markers exhibit trisomics segregation distortion in soybean [Glycine max (L.) Merr.]. Crop Sci. 46: 1456-1461.
Zou, J. J., R. J. Singh, and T. Hymowitz. 2004. SSR marker and ITS cleaved amplified polymorphic sequence analysis of soybean x Glycine tomentella intersubgeneric derived lines. Theor. Appl. Genet. 109:769-774.
Zou, J. J., R. J. Singh, J. Lee, S. J. Xu, P. B. Cregan, and T. Hymowitz. 2003. Assignment of molecular linkage groups to soybean chromosomes by primary trisomics. Theor. Appl. Genet. 107: 745-75.
Zou, J. J., R. J. Singh, and T. Hymowitz. 2003. Association of the yellow leaf-10 (y-10) mutant to soybean chromosome3. J. Hered. 94: 352-354.
Singh, R. J., H. H. Kim, and T. Hymowitz. 2001. Distribution of rDNA loci in the genus Glycine Willd. Theor. Appl. Genet. 103: 212-218.
Cregan, P. B., K. P. Kollipara, S. J. Xu, R. J. Singh, and T. Hymowitz.2001. Primary trisomics and SSR markers as tools to associate chromosomes with linkage groups in soybean. Crop Sci.41: 1262-1267.
Xu, S. J., R. J. Singh, K. P. Kollipara, and T. Hymowitz. 2000. Primary trisomics in soybean: origin, identification, breeding behavior, and use in gene mapping. Crop Sci. 40: 1543-1551.
Xu, S. J., R. J. Singh, and T. Hymowitz. 2000. Monosomics in soybean: origin, identification, cytology and breeding behavior. Crop Sci. 40: 985-989.
Xu, S. J., Singh, R. J., K. P. Kollipara, and T. Hymowitz. 2000. Hypertriploid in soybean: origin, identification, cytology and breeding behavior. Crop Sci. 40: 72-77.
Singh, R. J., and T. Hymowitz. 1999. Soybean genetic resources and crop improvement. Genome 42: 605-616.
Riggs, R. D., S. Wang, R. J. Singh, and T. Hymowitz. 1998. Possible transfer of resistance to Heterodera glycines from Glycine tomentella to Glycine max. J. Nematol. 30: 547-552.
Singh, R. J., K. P. Kollipara, and T. Hymowitz. 1998. Monosomic alien addition lines derived from Glycine max (L.) Merr. and G. tomentella Hayata: Production, characterization, and breeding behavior. Crop Sci.38: 1483-1489.
Singh, R. J., K. P. Kollipara, and T. Hymowitz. 1998. The genomes of Glycine canescens F. J. Herm., and G. tomentella Hayata of Western Australia and their phylogenetic relationships in the genus Glycine Willd. Genome 41: 669-679.
Singh, R. J., T. M. Klein, C. J. Mauvais, S. Knowlton, T. Hymowitz, and C. M. Kostow. 1998. Cytological characterization of the transgenic soybean. Theor. Appl. Genet. 96: 319 - 324.
Kollipara, K. P., R. J. Singh, and T. Hymowitz. 1997. Phylogenetic and genomic relationships in the genus Glycine Willd. based on sequence from the ITS region of nuclear rDNA. Genome 40: 57-68.
Singh, R. J., K. P. Kollipara, and T. Hymowitz. 1996. Development of hybrids between wild perennial soybeans and Glycine max (L.) Merr. and resistance to Phakopsora pachyrhizi Syd. In Soybean Rust Workshop, 9-11 August 1995. Edited by J.B. Sinclair and G.L. Hartman, College of Agriculture, Consumer, and Environmental Sciences, National Soybean Research Laboratory Publ. No.1, Urbana, Illinois. pp 46-51.
Kollipara, K. P., R. J. Singh, and T. Hymowitz. 1996. Inheritance of protease inhibitors in Glycine tomentella Hayata (2n=38), a perennial relative of soybean. J. Hered.87: 461-463
Kollipara, K. P., R. J. Singh, and T. Hymowitz. 1995. Genomic relationships in the genus Glycine (Fabaceae: Phaseoleae): Use of a monoclonal antibody to the soybean Bowman-Birk inhibitor as a genome marker. Amer. J. Bot. 82: 1104-1111.
Kollipara, K. P., R. J. Singh, and T. Hymowitz. 1994. Genomic diversity and multiple origins of tetraploid (2n=78, 80) Glycine tomentella Hayata. Genome 37: 448-459.
Kollipara, K. P., R. J. Singh, and T. Hymowitz. 1993. Genomic diversity in aneudiploid (2n=38) and diploid (2n=40) Glycine tomentella Hayata revealed by cytogenetic and biochemical methods. Genome 36: 391-396.
Singh, R. J., K. P. Kollipara, and T. Hymowitz. 1993. Backcross (BC2-BC4)-derived fertile plants from Glycine max (L.) Merr. and G. tomentella Hayata intersubgeneric hybrids. Crop Sci. 33: 1002-1007.
Singh, R. J., and T. Tsuchiya. 1993. Cytogenetics of an unstable trisomic in barley (Hordeum vulgare L.). Genome 36: 350-355.
Singh, R. J., and T. Tsuchiya. 1993. Cytogenetics of two novel compensating diploid in barley (Hordeum vulgare L.). Genome 36: 343-349.
Ahmad, F., R. J. Singh, and T. Hymowitz. 1992. Cytological evidence for four new primary trisomics in soybean. J. Hered. 83: 221-224.
Singh, R. J., K. P. Kollipara, and T. Hymowitz. 1992. Genomic relationships among diploid wild perennial species of the genus Glycine Willd. subgenus Glycine revealed by cytogenetics and seed protein electrophoresis. Theor. Appl. Genet. 85: 276-282.
Singh, R. J., K. P. Kollipara, F. Ahmad, and T. Hymowitz. 1992. Putative diploid ancestors of 80 - chromosome Glycine tabacina (Labill.) Benth. Genome 35: 140-146.
Singh, R. J., and T. Hymowitz. 1991. Cytogenetics of a synthesized allopentaploid (2n= 5x =100) in the genus Glycine Willd. subgenus Glycine. Genome 34: 751-756.
Griffor, M. C., L. O. Vodkin, R. J. Singh, and T. Hymowitz. 1991. Fluorescent in situ hybridization to soybean metaphase chromosomes. Plant Mol. Biol. 17: 101-109.
Singh, R. J., and F. L. Kolb. 1991. Chromosomal interchanges in six hexaploid oat genotypes. Crop Sci. 31: 726-729.
Singh, R. J., and T. Hymowitz. 1991. Identification of four primary trisomics of soybean by pachytene chromosome analysis. J. Hered. 82: 75-77.
Singh, R. J., K. P. Kollipara, and T. Hymowitz. 1990. Back cross-derived progeny from Soybean and Glycine tomentella Hayata intersubgeneric hybrids. Crop Sci. 30: 871-874.
Hymowitz, T., R. J. Singh, and R. P. Larkin. 1990. Long-distance dispersal: The case for the allopolyploids Glycine tabacina (Labill.) Benth. and G. tomentella Hayata in the West-Central Pacific. Micronesica 23: 5-13.
Singh, R. J., K. P. Kollipara, and T. Hymowitz. 1989. Ancestors of 80 and 78-chromosome Glycine tomentella Hayata (Leguminosae). Genome 32: 796-801.
Singh, R. J., and T. Hymowitz. 1989. The genomic relationships among Glycine soja Sieb. and Zucc., G. max (L.) Merr. and `G. gracilis' Skvortz. Plant Breed. 103: 171-173.
Singh, R. J., and T. Hymowitz. 1988. The genomic relationship between Glycine max (L.) Merr. and G. soja Sieb. and Zucc. as revealed by pachytene chromosome analysis. Theor. Appl. Genet. 76: 705-711.
Singh, R. J., K. P. Kollipara, and T. Hymowitz. 1988. Further data on the genomic relationships among wild perennial species (2n=40) of the genus Glycine Willd.. Genome 30: 166-176.
Singh, R. J., K. P. Kollipara, and T. Hymowitz. 1987. Polyploid complexes of Glycine tabacina (Labill.) Benth. and G. tomentella Hayata revealed by cytogenetic analysis. Genome 29: 490-497.
Singh, R. J., K. P. Kollipara, and T. Hymowitz. 1987. Intersubgeneric hybridization of soybeans with a wild perennial species, Glycine clandestina Wendl. Theor. Appl. Genet. 74: 391-396.
Singh, R. J., and T. Hymowitz. 1987. Intersubgeneric crossability in the genus Glycine Willd. Plant Breed. 98: 171-173.
Do Valle, C. B., R. J. Singh, and D. A. Miller. 1987. Pachytene chromosomes of Brachiaria ruziziensis Germain et Evrard. Plant Breed. 98: 75-78.
Singh, R. J. 1986. Chromosomal variation in immature embryo derived calluses of barley (Hordeum vulgare L.). Theor. Appl. Genet. 72: 710-716.
Singh, R. J., and T. Hymowitz. 1985. Diploid-like meiotic behavior in synthesized amphiploids of the genus Glycine Willd. subgenus Glycine. Can. J. Genet. Cytol. 27: 655-660.
Singh, R. J., and T. Hymowitz. 1985. The genomic relationships among six wild perennial species of the genus Glycine subgenus Glycine Willd. Theor. Appl. Genet. 71: 221-230.
Singh, R. J., and T. Hymowitz. 1985. Intra- and interspecific hybridization in the genus Glycine subgenus Glycine Willd.: Chromosome pairing and genome relationships. Z. Pflanzenzüchtg. 95: 289-310
Singh, R. J., and T. Hymowitz. 1985. An intersubgeneric hybrid between Glycine tomentella Hayata and the soybean, G. max (L.) Merr. Euphytica 34: 187-192.
Tsuchiya, T., R. J. Singh, A. Shahla, and A. Hang. 1984. Acrotrisomic analysis in linkage mapping in barley (Hordeum vulgare L.) Theor. Appl. Genet. 68: 433-440.
Singh, R. J. 1984 The regeneration of milkweed (Asclepias spp.) plants from hypocotyl callus, cell suspension cultures and hypocotyl protoplast. Z. Pflanzenzüchtg. 92: 95-101.
Khush, G. S., R. J. Singh, S. C. Sur, and A. L. Librojo. 1984. Primary trisomics of rice: Origin, morphology, cytology and use in linkage mapping. Genetics 107: 141-163.
Singh, R. J., G. S. Khush, and T. W. Mew. 1983. A new gene for resistance to bacterial blight in rice. Crop Sci. 23: 558-560.
Singh, R. J., and T. Tsuchiya. 1982. Identification and designation of telocentric chromosomes in barley by means of Giemsa N-banding technique. Theor. Appl. Genet. 64: 13-24.
Tsuchiya, T., and R. J. Singh. 1982. Chromosome mapping in barley by means of telotrisomic analysis. Theor. Appl. Genet. 61: 201-208.
Singh, R. J., and T. Tsuchiya. 1982. An improved Giemsa N-banding technique for the identification of barley chromosomes. J. Hered. 73: 227-229.
Singh, R. J., and T. Tsuchiya. 1981. Identification and designation of barley chromosomes by Giemsa banding technique: a reconsideration. Z. Pflanzenzüchtg 86: 336-340.
Singh, R. J., and T. Tsuchiya. 1981. Cytological study of the telocentric chromosomes in seven monotelotrisomics of barley. Bot. Gaz. 142: 267-273.
Singh, R. J., and T. Tsuchiya. 1981. Ring chromosomes in barley (Hordeum vulgare L.). Chromosoma (Berl.) 82: 133-141.
Singh, R. J., and H. Ikehashi. 1981. Monogenic male sterility in rice: Induction, identification and inheritance. Crop Sci. 21: 286-289.
Singh, R. J., and T. Tsuchiya. 1977. Morphology, fertility, and transmission in seven monotelotrisomics of barley. Z. Pflanzenzüchtg. 78: 327-340.
Singh, R. J. 1977. Evidence of heterochromatin polymorphism through crossing over. Wheat Inf. Serv. 44: 19-21.
Singh, R. J. 1977. Cross compatibility, meiotic pairing and fertility in five Secale species and their interspecific hybrids. Cereal Res. Comm. 5: 67-75.
Singh, R. J., and G. Röbbelen. 1977. Identification by Giemsa technique of the translocation separating cultivated rye from three wild species of Secale. Chromosoma (Berl.) 59: 217-225.
Singh, R. J., G. Röbbelen, and M. Okamoto. 1976. Somatic association at interphase studied by Giemsa banding technique. Chromosoma (Berl.) 56: 265-273.
Singh, R. J., and G. Röbbelen. 1976. Giemsa banding technique reveals deletions within rye chromosomes in addition lines. Z. Pflanzenzüchtg. 76: 11-18.
Singh, R. J., and G. Röbbelen. 1975. Comparison of somatic Giemsa banding pattern in several species of rye. Z. Pflanzenzüuchtg. 75: 270-285.
Singh, R. J., and T. Lelley. 1975. Giemsa banding in meiotic chromosomes of rye, Secale cereale L. Z. Pflanzenzüchtg. 75: 85-89.
Singh, R. J., and T. Tsuchiya. 1975. Pachytene chromosomes of barley. J. Hered. 66: 165-167.
Singh, R. J., and T. Tsuchiya. 1975. Hypertriploid plants in barley (Hordeum vulgare L.). Caryologia 28: 89-98.
Singh, R. J. 1970. Hybrid vigor in Bajra (Pennisetum typhoides S. & H.). Indian J. Agr. Sci. 40: 974-976.
Gill, N. S., and R. J. Singh. 1970. Effect of soil substrate and re treatment on cold test response of seed corn (Zea mays L.). Proc. Assoc. Off. Seed Anal. 60:181-187.
Books
Genetic Resources Chromosome Engineering and Crop Improvement Volume 6 Medicinal Plants (Pub. 2011)
Genetic Resources Chromosome Engineering and Crop Improvement Volume 5 Forage Crops (Pub. 2009)
Genetic Resources Chromosome Engineering and Crop Improvement Volume 4 Oilseed Crops (Pub. 2007)
Genetic Resources Chromosome Engineering and Crop Improvement Volume 3 Vegetable Crops (Pub. 2007)
Genetic Resources Chromosome Engineering and Crop Improvement Volume 2 Cereals (Pub. 2006)
Genetic Resources Chromosome Engineering and Crop Improvement Volume 1 Grain Legumes (Pub. 2005)
Plant Cytogenetics First Edition (Pub. 1993)
Plant Cytogenetics Second Edition (Pub. 2003)
Contributed Chapters
Singh, R. J., A. Lebeda, and, A. O. Tucker. 2011. Medicinal plants-natures pharmacy. In: Genetic Resources, Chromosome Engineering, and Crop Improvement Series: Medicinal Plants- Volume 6. Edited by Ram J. Singh, CRC Press (Taylor & Francis Group). pp 13-51
Singh, R. J. 2011. Landmark research in medicinal plants. In: Genetic Resources, Chromosome Engineering, and Crop Improvement Series: Medicinal Plants- Volume 6. Edited by Ram J. Singh, CRC Press (Taylor & Francis Group). pp. 1-11
Ratnaparkhe, M. B., R. J. Singh, and J. J. Doyle. 2011. Glycine In: Wild Crop Relatives: Genomics and Breeding Resources, Legumes Crops and Forages. Edited by C. Kole, Springer-Verlag Berlin Heidelberg. pp. 83-116.
Singh R. J., and W. F. Grant. 2009. Landmark Research in Forage Crops. In: Genetic Resources, Chromosome Engineering, and Crop Improvement Series: Forage Crops- Volume 5. Edited by Ram J. Singh CRC Press (Taylor & Francis Group). pp. 1-9
Singh, R. J., R. L. Nelson, and G. H. Chung. 2007. Soybean [Glycine max (L.) Merr.]. In: Genetic Resources, Chromosome Engineering, and Crop Improvement Series: Oilseed Crops- Volume 4 . Edited by Ram J. Singh CRC Press (Taylor & Francis Group). pp. 13-50.
Singh, R. J., 2007. Landmark Research in Oilseed Crops. In: Genetic Resources, Chromosome Engineering, and Crop Improvement Series: Oilseed Crops- Volume 4 . Edited by Ram J. Singh CRC Press (Taylor & Francis Group). pp. 1-12.
Singh, R. J., and A. Lebeda. 2007. Landmark Research in Vegetable Crops. In: Genetic Resources, Chromosome Engineering, and Crop Improvement Series: Vegetable Crops- Volume 3 . Edited by Ram J. Singh, CRC Press (Taylor & Francis Group). pp. 1-15
Costa, J. M., and R. J. Singh. 2005. Chromosome Mapping in Barley (Hordeum vulgare L.). In: Genetic Resources, Chromosome Engineering, and Crop Improvement Series: Cereals-Volume 2. Edited by Ram J. Singh, and Prem P. Jauhar. CRC Press (Taylor & Francis Group). pp. 257-280.
Singh, R. J. 2005. Utilization of Genetic Resources for Barley Improvement. In: Genetic Resources, Chromosome Engineering, and Crop Improvement Series: Cereals- Volume 2. Edited by Ram J. Singh, and Prem P. Jauhar. CRC Press (Taylor & Francis Group. pp. 233-255
Singh, R. J. 2005. Landmark Research in Grain Legumes. In: Genetic Resources, Chromosome Engineering, and Crop Improvement Series: Grain Legumes- Volume 1. Edited by Ram J. Singh, and Prem P. Jauhar. CRC Press (Taylor & Francis Group). pp. 1-9.
Singh, R. J. 2002. Chromosomal and Genetic Aberrations in Transgenic Soybean. In: Molecular Methods of Plant Analysis: “Testing for Genetic manipulation”. Edited by J. F. Jackson, H. F. Linskens and R. B. Inman. Springer-Verlag Berlin Heidelberg. pp. 153-168.
Singh, R. J., and G. S. Khush. 2000. Cytogenetics of Rice. In: Rice Breeding and Genetics- Research Priorities and Challanges. Edited by J. S. Nanda, Science Pub. Inc., Enfield, New Hampshire, U.S.A. pp. 287-311.
Hymowitz, T., R. J. Singh, and K. P. Kollipara, 1998. The Genomes of the Genus Glycine. Plant Breed Rev.16: 289-317.
Hymowitz, T., R. G. Palmer, and R. J. Singh. 1991. Cytogenetics of the Genus Glycine. In Chromosome Engineering in Plants: Genetics, Breeding and Evolution, Edited by T. Tsuchiya and P. K. Gupta, Elsevier Sci. Pub., Amsterdam, The Netherlands. pp. 53-63.
Khush, G. S., and R. J. Singh. 1991. Chromosome Architecture and Aneuploidy in Rice. In Chromosome Engineering in Plants: Genetics, Breeding and Evolution, Edited by T. Tsuchiya and P. K. Gupta, Elsvier Sci. Pub., Amsterdam, The Netherlands. pp. 577-598.
Hymowitz, T., and R. J. Singh. 1987. Taxonomy and Speciation. In Soybeans: Improvement, Production and Uses - Revision, Edited by J. R. Wilcox, ASA, Madison, Wisconsin. pp. 23-48.
Khush, G. S. and R. J. Singh. 1986. Relationships between Linkage Groups and Cytologically Identifiable Chromosomes of Rice. In Rice Genetics. Island Publishing House Inc. Manila, Philippines. pp. 239-248.
Patents
Singh, R. J. 2010. Methods for producing fertile crosses between wild and domestic soybean species. Patent No.: US 7,842,850 (Date of patent Nov. 30, 2010)